Research article

Analysis of a stochastic two-species Schoener's competitive model with Lévy jumps and Ornstein–Uhlenbeck process

  • Received: 27 December 2023 Revised: 13 March 2024 Accepted: 21 March 2024 Published: 28 March 2024
  • MSC : 60H10, 60H30, 92D25

  • This paper studies a stochastic two-species Schoener's competitive model with Lévy jumps by the mean-reverting Ornstein–Uhlenbeck process. First, the biological implication of introducing the Ornstein–Uhlenbeck process is illustrated. After that, we show the existence and uniqueness of the global solution. Moment estimates for the global solution of the stochastic model are then given. Moreover, by constructing the Lyapunov function and applying Itô's formula and Chebyshev's inequality, it is found that the model is stochastic and ultimately bounded. In addition, we give sufficient conditions for the extinction of species. Finally, numerical simulations are employed to demonstrate the analytical results.

    Citation: Yajun Song, Ruyue Hu, Yifan Wu, Xiaohui Ai. Analysis of a stochastic two-species Schoener's competitive model with Lévy jumps and Ornstein–Uhlenbeck process[J]. AIMS Mathematics, 2024, 9(5): 12239-12258. doi: 10.3934/math.2024598

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  • This paper studies a stochastic two-species Schoener's competitive model with Lévy jumps by the mean-reverting Ornstein–Uhlenbeck process. First, the biological implication of introducing the Ornstein–Uhlenbeck process is illustrated. After that, we show the existence and uniqueness of the global solution. Moment estimates for the global solution of the stochastic model are then given. Moreover, by constructing the Lyapunov function and applying Itô's formula and Chebyshev's inequality, it is found that the model is stochastic and ultimately bounded. In addition, we give sufficient conditions for the extinction of species. Finally, numerical simulations are employed to demonstrate the analytical results.



    As a common form of ecosystem, competition relationship has been widely applied in many fields in recent years, and many scholars have conducted detailed research on the competition relationship between populations [1]. In 2005, Zhen et al. [2] studied a class of two-dimensional non-autonomous competitive Lotka-Volterra systems with pulses. In 2012, Liu et al. [3] investigated the quasi-periodic solution of a Lotka-Volterra competitive system with quasi-periodic perturbations. In 2014, Liu et al. [4] studied stochastic logistic models with Lévy noise and obtained necessary and sufficient conditions for stochastic persistence and extinction. The Lotka-Volterra competition model is widely employed to study a variety of systems, including prey-predator dynamics in ecological interaction[5,6,7]. However, its linearization poses limitations to modeling the dynamics of certain species[8,9], for which more complex approaches are required. In 1974, a more practical competitive system was proposed and discussed by Schoener [10], as follows :

    {dx(t)=x(t)[r1β1x(t)α1y(t)+ξ1x(t)+δ1]dt,dy(t)=y(t)[r2β2x(t)α2y(t)+ξ2y(t)+δ2]dt, (1.1)

    where x(t) and y(t) reflect the population density at time t, r1 and r2 represent the death rates of the population, β1 and α2 represent the intra-specific competition rates, and β2 and α1 represent the inter-specific competition rates. All parameters in system (1.1) are assumed to be positive constants. In recent years, more and more scholars have studied Schoener's competition model and achieved many results. In 2008, Wu et al.[11] studied a class of discrete Schoener's competition models with time delay, and obtained sufficient conditions to ensure the durability of the model and the global attraction of the model's positive solution. In 2012, LV et al.[8] investigated the dynamics of Schoener's competition model for two stochastic populations. In 2016, Li et al.[10] obtained some sufficient conditions for the persistence of uniformly asymptotically stable positive almost periodic solutions for a class of pulsed Schoener competition model with pure-delays.

    However, in the natural world, population systems are inevitably affected by environmental noise. Therefore, considering only deterministic models can be very flawed. Many parameters in ecological dynamics should fluctuate around some average values. Mao et al. [12] demonstrated that even small amounts of ambient noise can have a large impact on species populations, which means that stochastic population models can provide additional authenticity compared to deterministic population models. Therefore, it is generally assumed that environmental noise primarily affects the fundamental parameters of the model in order to study the dynamic properties of ecosystems in different environments[13]. Based on the fact that population death rates are easily affected by environmental fluctuations, we assume r1 and r2 in Schoener's competition model are two random variables. Currently, there are two common methods for simulating small disturbances in the environment. The most common method is to introduce Gaussian linear white noise into the deterministic model[14,15,16,17]. Another method is to incorporate the mean-reverting Ornstein-Uhlenbeck process to simulate environmental perturbations[18,19,20], which has been demonstrated to be a practical and biologically meaningful method.

    First, we assume that the death rate is linearly correlated with Gaussian white noise in the random environment. That is to say,

    ri(t)=ri+σidBi(t)dt,i=1,2,

    where ri represents the long-time average levels of ri(t). Bi(t) denotes two independent standard Brownian motions defined on a complete probability space {Ω,F,{Ft}t0,P} with a filtration {Ft}t0 satisfying the usual conditions[12]. For any time interval [0,t], we can obtain that

    ri(t):=1tt0ri(s)ds=ri+σiBi(t)tN(ri,σ2it),i=1,2,

    where ri(t) is the time average of ri(t), and N(,) denotes the one-dimensional normal distribution. From the above formula, it is easy to see that the variance of ri(t) is σ2it, which tends to infinity as t0+. This means that the mean value of the perturbation parameter will be more variable in a small amount of time. In practice, however, ri(t) should likewise fluctuate little over a small time interval. Thus, the use of Gaussian linear white noise to simulate small disturbances in the environment is unreasonable.

    We now consider the second method, which is to introduce the Ornstein-Uhlenbeck process into the deterministic model. On account of this approach, one has

    dr1(t)=ω1(r1r1)dt+σ1dB1(t),dr2(t)=ω2(r2r2)dt+σ2dB2(t), (1.2)

    where ωi>0 and σi>0 are the speed of reversion and the intensity of volatility, respectively. By performing a stochastic integral operation on Equation (1.2), we can get the following unique explicit solutions:

    ri(t)=ri+[ri(0)ri]eωit+σit0eωi(ts)dBi(s),i=1,2, (1.3)

    where ri(0) is the initial value of the Ornstein–Uhlenbeck process ri(t). For any time interval [0,t], we have

    ri(t):=1tt0ri(s)ds=ˉri+1tt0σiωi(1eωi(st))dBi(s)N(ˉri,σ2it3+O(t2)),

    and as the time interval t0+, it is clear that E[ri(t)]=ˉri and Var[ri(t)]=0, which is consistent with the fact that the perturbation of the death rate in a small time interval is also small. Therefore, it is more reasonable to introduce the Ornstein-Uhlenbeck process to perturb the parameters ri(t),i=1,2, than Gaussian white noise[20,21].

    Based on the above analysis, we decided to introduce the Ornstein-Uhlenbeck process into the studied model. By combining system (1.1) and system (1.2), we can obtain a stochastic model of the following form:

    {dx(t)=x(t)[r1(t)β1x(t)α1y(t)+ξ1x(t)+δ1]dt,dy(t)=y(t)[r2(t)β2x(t)α2y(t)+ξ2y(t)+δ2]dt,dr1(t)=ω1[ˉr1r1(t)]dt+σ1dB1(t),dr2(t)=ω2[ˉr2r2(t)]dt+σ2dB2(t). (1.4)

    However, in addition to small perturbations in the environment, population dynamics can be subject to sudden and violent environmental shocks, such as avalanches, earthquakes, and tsunamis. It is important to note that these environmental shocks may cause sharp jumps in the population size so that previously continuous solution trajectories are no longer continuous. However, this phenomenon cannot be described by Brownian motion, so stochastic differential equations with Lévy jumps are often used to simulate sudden random perturbations that occur in the environment[22,23]. In 2014, Liu et al. [24] studied the Lotka-Volterra stochastic model disturbed by Lévy noise and established the necessary and sufficient conditions for persistence in the mean and extinction of each population. In 2018, Qiu and Deng[1] studied the optimal acquisition problem of randomly competing Lotka-Volterra models under the influence of time delay. Inspired by the above, this paper adds Lévy jumps to system (1.4) to simulate sudden environmental disturbances in nature. It takes the form

    {dx(t)=x(t)[r1(t)β1x(t)α1y(t)+ξ1x(t)+δ1]dt+Yx(t)γ1(μ)N(dt,dμ),dy(t)=y(t)[r2(t)β2x(t)α2y(t)+ξ2y(t)+δ2]dt+Yy(t)γ2(μ)N(dt,dμ),dr1(t)=ω1[ˉr1r1(t)]dt+σ1dB1(t),dr2(t)=ω2[ˉr2r2(t)]dt+σ2dB2(t), (1.5)

    where x(t), y(t), and r(t) represent the left limit at t, N(,) is a Poisson counting measure with characteristic measure λ on a measurable subset Y of (0,+) with λ(Y)<, and N(,) is independent of Brownian motion B(). Define the compensating random measure as ˜N, then N(dt,dμ)=˜N(dt,dμ)+λ(dμ)dt. γ() represents the disturbance intensity of Lévy jump noise to the population.

    The model established in this paper is an improvement on the classical Schoener's competition model. First, system (1.5) uses the mean-reverting Ornstein-Uhlenbeck process to simulate small perturbations in the environment, which is more reasonable than assuming that population parameters are linearly distributed in Gaussian white noise. In addition, considering that species in the real world are often affected by sudden random perturbations, we also introduced Lévy jumps into the model. Therefore, this model takes into account both small environmental disturbances and sudden violent disturbances, which can better simulate some random phenomena in nature.

    The rest of the paper is organized as follows. Section 2 gives some necessary lemmas and assumptions. Section 3 shows several dynamical properties of system (1.5), including the existence and uniqueness of global solutions, moment estimate, and stochastic ultimate boundness. In addition, we obtain sufficient conditions for species extinction. In Section 4, we carry out some numerical simulations to verify the theoretical results. Finally, we give some conclusions in Section 5.

    Remark 1. The model studied in this paper perturbs the parameter ri, where ri represents the death rate of the population. Due to the characteristics of the Ornstein-Uhlenbeck process, the death rate can be taken as any real number in the model. However, it is well known that negative death rates are biologically implausible. For this phenomenon, we have the following explanation:

    First, the positive or negative death rate does not affect the proof of the theorem, so even if the death rate may be negative, it does not affect the results obtained in this paper. Moreover, since the theorem is true for any value of death rate, it is also true for non-negative death rate. It is worth noting that the negative death rate is only a theoretical value, and we still assume that it is positive in practice and numerical simulation, that is, we mainly use the part of ri0,i=1,2.

    Due to the need of subsequent proofs, several lemmas and assumptions will be given in this section.

    Lemma 2.1. (Chebyshev inequality) If c>0, p>0, XLp, we have

    P{ω:|X(ω)|c|}cpE[|X|p],

    where Lp represents the set of random variables that take values on Rn and E[|X|p]<.

    Lemma 2.2. Let M(t) be a local martingale with initial value M(0)=0. If limtρM(t)<, then

    limtM(t)t=0,

    where

    ρM(t)=t0dM,M(s)(1+s)2, t0.

    Assumption 2.1. For any i{1,2}, there is always a constant c that makes the following true:

    (1) Y(|ln(1+γi(μ))||ln(1+γi(μ))|2)λ(μ)<c.(2) Y|γi(μ)|λ(dμ)<c.(3) Y|(1+γi(μ))q1|λ(dμ)<c. (2.1)

    Theorem 3.1. For any initial value condition

    (x(0),y(0),r1(0),r2(0))R2+×R2,

    system (1.5) has a unique solution (x(t),y(t),r1(t),r2(t)) on t0, and it will remain in R2+×R2 with probability one.

    Proof. For t0, and for any initial value (x(0),y(0),r1(0),r2(0))R2+×R2, it is easy to prove that the equation coefficients in system (1.5) satisfy local Lipschitz conditions. Therefore, system (1.5) has a unique local solution (x(t),y(t),r1(t),r2(t))R2+×R2 on t[0,τe), where τe is the explosion time[12].

    To prove that the model has a global positive solution, we only need to show that τe= a.s.. By defining a necessary set Hn0=(n0,n0)×(n0,n0)×(n0,n0)×(n0,n0), we can always determine a sufficiently large integer n0 such that (lnx(0),lny(0),r1(0),r2(0))Hn0. For any integer nn0, we define a stopping time set τn by

    τn=inf{t[0,τe)|lnx(t)(n,n),or lny(t)(n,n),or r1(t)(n,n),or r2(t)(n,n)}. (3.1)

    Obviously, τn is monotonically increasing as n increases. For convenience, let τ=limnτn and inf=, which implies ττe a.s.. To prove Theorem 3.1, it suffices to verify τ=  a.s.. Consider the contradiction, i.e., τ< a.s.. Then there are constants T>0 and ε(0,1) to make P{τT}>ε. Hence, there is a positive number n1n0 such that

    P{τnT}ε,nn1. (3.2)

    For any tτn, by the inequality x1lnx (x>0), a non-negative C2-function V(x(t),y(t),r1(t),r2(t)) is constructed as follows:

    V(x,y,r1,r2)=x(t)1lnx(t)+y(t)1lny(t)+r41(t)4+r42(t)4. (3.3)

    Applying the Itˆo formula yields,

    dV=LVdt+r31(t)σ1dB1(t)+r32(t)σ2dB2(t)+Y(x(t)γ1(μ)ln(1+γ1(μ)))˜N(dt,dμ)+Y(y(t)γ2(μ)ln(1+γ2(μ)))˜N(dt,dμ), (3.4)

    where

    LV=x(t)r1(t)β1x2(t)α1x(t)y(t)+ξ1x(t)x(t)+δ1+r1(t)+β1x(t)+α1y(t)ξ1x(t)+δ1y(t)r2(t)β2y(t)x(t)α2y2(t)+ξ2y(t)y(t)+δ2+r2(t)+β2x(t)+α2y(t)ξ2y(t)+δ2+ω1r31(t)(ˉr1r1(t))+ω2r32(t)(ˉr2r2(t))+32σ21r21(t)+32σ22r22(t)+Y(x(t)γ1(μ)ln(1+γ1(μ)))λ(dμ)+Y(y(t)γ2(μ)ln(1+γ2(μ)))λ(dμ). (3.5)

    Combined with Assumption 2.1, it is easy to know that there exists an upper bound K such that

    LV|r1(t)|(1+x(t))+ξ1x(t)x(t)+δ1+(β1+β2)x(t)+|r2(t)|(1+y(t))+ξ2y(t)y(t)+δ2+(α1+α2)y(t)+ω1ˉr1r31(t)+ω2ˉr2r32(t)+32σ21r21(t)+32σ22r22(t)β1x2(t)α2y2(t)(α1+β2)x(t)y(t)+x(t)Yγ1(μ)λ(dμ)+y(t)Yγ2(μ)λ(dμ)ξ1x(t)+δ1ξ2y(t)+δ2ω1r41(t)ω2r42(t)+2cK,

    thus

    dVKdt+r31(t)σ1dB1(t)+r32(t)σ2dB2(t)+Y(x(t)γ1(μ)ln(1+γ1(μ)))˜N(dt,dμ)+Y(y(t)γ2(μ)ln(1+γ2(μ)))˜N(dt,dμ). (3.6)

    Integrating the inequality from 0 to τnT and then taking the expectation on both sides of inequality (3.7), we have

    E[V(x(τnT),y(τnT),r1(τnT),r2(τnT))]V(x(0),y(0),r1(0),r2(0))+KT. (3.7)

    For all n1n0, let Ωn={τnT}. Then we have P(Ωn)ε. Note that, for any ωΩn, x, y, r1, and r2 equals either n or n, so there is

    V(x(τn,ω),y(τn,ω),r1(τn,ω),r2(τn,ω))min{en1+n,en1n}.

    According to the inequality 3.7, it can be derived that

    V(x(0),y(0),r1(0),r2(0))+KTE[1Ωn(ω)V(x(τn),y(τn),r1(τn),r2(τn))]εmin{en1+n,en1n}, (3.8)

    where 1Ωn(ω) represents the index function. As n, we have >V(x(0),y(0),r1(0),r2(0))+KT, which leads to a contradiction. Therefore, we have τ= a.s.. This completes the proof of Theorem 3.1.

    In this section, we will provide a moment estimate for the global solution mentioned above.

    Theorem 3.2. For any initial value (x(0),y(0),r1(0),r2(0))R2+×R2, the solution (x(t),y(t),r1(t),r2(t)) of system (1.5) has the property

    E[|x(t),y(t)|q]K(q) (3.9)

    for any q>0, where K(q) is a continuous function with respect to q. That is to say, the qth moment of the solution x(t),y(t) is bounded.

    Proof. For any q2, we define a non-negative C2-function

    V1(x(t),y(t),r1(t),r2(t)):R2+×R2

    by

    V1(x(t),y(t),r1(t),r2(t))=xq(t)q+yq(t)q+r2q1(t)2q+r2q2(t)2q.

    Applying the generalized Itˆo formula, we obtain

    dV1=LV1dt+r2q11(t)σ1dB1(t)+r2q12(t)σ2dB2(t)+xq(t)qY((1+γ1(μ))q1)˜N(dt,dμ)+yq(t)qY((1+γ2(μ))q1)˜N(dt,dμ). (3.10)

    To simplify the notation, the subsequent proof process replaces x(t), y(t), ri(t), and γi(μ) with x, y, ri, and γi, respectively. So, we have

    LV1=xqr1β1xq+1α1xqy+ξ1xqx+δ1yqr2β2xyqα2yq+1+ξ2yqy+δ2+ω1r2q11(ˉr1r1)+ω2r2q12(ˉr2r2)+2q12σ21r2q21+2q12σ22r2q22+xqqY((1+γ1)q1)λ(dμ)+yqqY((1+γ2)q1)λ(dμ). (3.11)

    Taking the mathematical expectation of eηtV1, we obtain

    E(eηtV1(x(t),y(t),r1(t),r2(t)))=E(V1(x(0),y(0),r1(0),r2(0)))   +t0E[L(eηsV1(x(s),y(s),r1(s),r2(s)))]ds, (3.12)

    where η=qmin{ω1,ω2}. Noting that

    L[eηtV1(x,y,r1,r2)]=ηeηtV1(x,y,r1,r2)+eηtLV1(x,y,r1,r2)=eηt{ηqxq+ηqyqxqr1β1xq+1α1xqy+ξ1xqx+δ1yqr2β2xyqα2yq+1+ξ2yqy+δ2(ω1η2q)r2q1(ω2η2q)r2q2+ω1r2q11ˉr1+ω2r2q12ˉr2+2q12σ21r2q21+2q12σ22r2q22+xqqY((1+γ1)q1)λ(dμ)+yqqY((1+γ2)q1)λ(dμ)}eηt{ω1xq+ω2yq+|r1|xq+|r2|yq+ξ1xqx+δ1+ξ2yqy+δ2+ω1r2q11ˉr1+ω2r2q12ˉr2+2q12σ21r2q21+2q12σ22r2q22β1xq+1α2yq+1α1xqyβ2xyqω12r2q1ω22r2q2+xqqY((1+γ1)q1)λ(dμ)+yqqY((1+γ2)q1)λ(dμ)}k(q)eηt, (3.13)

    where

    k(q)=sup(x,y,r1,r2)R2+×R2{ω1xq+ω2yq+|r1|xq+|r2|yq+ξ1xqx+δ1+ξ2yqy+δ2+ω1r2q11ˉr1+ω2r2q12ˉr2+2q12σ21r2q21+2q12σ22r2q22β1xq+1α2yq+1α1xqyβ2xyqω12r2q1ω22r2q2+xqqY((1+γ1)q1)λ(dμ)+yqqY((1+γ2)q1)λ(dμ)}<.

    Combining formula (3.12) and formula (3.13), we obtain

    E(eηtV1(x(t),y(t),r1(t),r2(t)))E(V1(x(0),y(0),r1(0),r2(0)))+k(q)(eηt1)η, (3.14)

    and then we have

    lim suptE[|x(t),y(t)|q]qlim suptE[V1(x(t),y(t),r1(t),r2(t))]limtE[V1(x(0),y(0),r1(0),r2(0))]eηt+limtk(q)(eηt1)ηeηt=0+k(q)η=k(q)ηa.s.. (3.15)

    The continuity of x(t),y(t) on t[0,) along with this implies that there exists a constant K(q) such that

    E[|x(t),y(t)|q]K(q),  t0,q2.

    By applying H¨older's inequality, we can conclude that, for any q(0,2),

    E(|x(t),y(t)|q)(E(|x(t),y(t)|)2)q2(K(2))q2.

    This completes the proof of Theorem 3.2.

    First, the definition of stochastic ultimate boundedness is given.

    Definition 3.1. The solution of system (1.5) is said to be stochastic and ultimately bounded, if for any ε(0,1), there is a positive number H(=H(ε)) such that for any initial value (x(0),y(0),r1(0),r2(0))R2+×R2, the solution of the model satisfies

    lim suptP{x2(s)+y2(s)H}1ε.

    Theorem 3.3. The solutions of system (1.5) are stochastic and ultimately bounded.

    Proof. According to Theorem 3.2, the solution of system (1.5) is moment bounded. Let q=0.5, then there is a positive number K0 such that for any initial value (x(0),y(0),r1(0),r2(0))R2+×R2, the solution of system (1.5) satisfies

    lim suptE(x2(s)+y2(s))12K0.

    According to the Lemma 2.1: P{|x(t)|>H}E|x(t)|H, by setting H=K0/ε, we can get

    lim suptP{x2(s)+y2(s)>H}ε,

    then

    lim suptP{x2(s)+y2(s)H}1ε.

    This completes the proof of Theorem 3.3.

    Theorem 3.4. For any initial value (x(0),y(0),r1(0),r2(0))R2+×R2, the solution (x(t),y(t),r1(t),r2(t)) of system (1.5) has the property that

    lim suptlnx(t)tlim supt1tt0(r1+ξ1δ1+Yln(1+γ1)λ(dμ))ds,
    lim suptlnx(t)tlim supt1tt0(r2+ξ2δ2+Yln(1+γ2)λ(dμ))ds.

    In particular, if

    G1=lim supt1tt0(r1+ξ1δ1+Yln(1+γ1)λ(dμ))ds<0,
    G2=lim supt1tt0(r2+ξ2δ2+Yln(1+γ2)λ(dμ))ds<0,

    then x(t), y(t) are extinct.

    Proof. Applying the Itˆo formula to lnx(t) and lny(t), it can be obtained that

    dlnx(t)=(r1β1xα1y+ξ1x+δ1+Yln(1+γ1)λ(dμ))dt+Yln(1+γ1)˜N(dt,dμ), (3.16)
    dlny(t)=(r2β2xα2y+ξ2y+δ2+Yln(1+γ2)λ(dμ))dt+Yln(1+γ2)˜N(dt,dμ). (3.17)

    Integrating from 0 to t, we have

    lnx(t)=lnx(0)+t0(r1β1xα1y+ξ1x+δ1+Yln(1+γ1)λ(dμ))ds+t0Yln(1+γ1)˜N(ds,dμ), (3.18)
    lny(t)=lny(0)+t0(r2β2xα2y+ξ2y+δ2+Yln(1+γ2)λ(dμ))ds+t0Yln(1+γ2)˜N(ds,dμ). (3.19)

    According to equalities (3.18) and (3.19), it can be obtained that

    lnx(t)lnx(0)+t0(r1+ξ1δ1+Yln(1+γ1)λ(dμ))ds+t0Yln(1+γ1)˜N(ds,dμ), (3.20)
    lny(t)lny(0)+t0(r2+ξ2δ2+Yln(1+γ2)λ(dμ))ds+t0Yln(1+γ2)˜N(ds,dμ). (3.21)

    Let ˜M1=t0Yln(1+γ1)˜N(dt,dμ), ˜M2=t0Yln(1+γ2)˜N(dt,dμ). According to Lemma 2.2, we have

    limt˜M1(t)t=0,limt˜M2(t)t=0.

    Then,

    lim suptlnx(t)tlim supt1tt0(r1+ξ1δ1+Yln(1+γ1)λ(dμ))ds, (3.22)
    lim suptlny(t)tlim supt1tt0(r2+ξ2δ2+Yln(1+γ2)λ(dμ))ds. (3.23)

    Therefore, when

    G1=lim supt1tt0(r1+ξ1δ1+Yln(1+γ1)λ(dμ))ds<0,
    G2=lim supt1tt0(r2+ξ2δ2+Yln(1+γ2)λ(dμ))ds<0,

    it implies limtx(t)=0, limty(t)=0, and then x(t), y(t) are extinct. This completes the proof of Theorem 3.4.

    In this section, we will verify the theoretical results by numerical simulation examples. First, the system (1.5) is discretized using the Milstein scheme for higher order discretization[25]. Consider the following stochastic Schoener's competitive model for two populations:

    {dx(t)=x(t)[r1(t)β1x(t)α1y(t)+ξ1x(t)+δ1]dt+Yx(t)γ1(μ)N(dt,dμ),dy(t)=y(t)[r2(t)β2x(t)α2y(t)+ξ2y(t)+δ2]dt+Yy(t)γ2(μ)N(dt,dμ),dr1(t)=ω1[ˉr1r1(t)]dt+σ1dB1(t),dr2(t)=ω2[ˉr2r2(t)]dt+σ2dB2(t). (4.1)

    Immediately afterwards, it is necessary to introduce the biological significance of the parameters related to the process being modeled. Computer simulations can then be performed to gain insights into the dynamics of the biological system (see Table 1).

    Table 1.  List of biological parameters.
    Parameter Explication
    ˉr1 Average mortality of species x
    ˉr2 Average mortality of species y
    β1 The intra-specific competition rates of species x
    β2 The inter-specific competition rates
    α1 The inter-specific competition rates
    α2 The intra-specific competition rates of species y
    σ1 The intensity of volatility of r1
    σ2 The intensity of volatility of r2

     | Show Table
    DownLoad: CSV

    Based on the biological significance of the above parameters, and in combination with reference[8,9,10,26], we choose the reasonable values as shown in Table 2.

    Table 2.  Several combinations of biological parameters of system (1.5) in Table 1.
    Combinations Value
    (A1)[8] ˉr1=0.15, ˉr2=0.2, β1=0.4, β2=0.1, α1=0.1, α2=0.1, ξ1=0.3, ξ2=0.2, δ1=0.6
    δ2=0.1, γ1=0.5, γ2=0.5, ω1=0.1, ω2=0.1, σ1=0.04, σ2=0.04
    (A2)[9] ˉr1=0.4, ˉr2=0.2, β1=0.3, β2=0.4, α1=0.4, α2=0.2, ξ1=0.65, ξ2=0.5, δ1=0.65
    δ2=0.55, γ1=0.2, γ2=0.2, ω1=0.1, ω2=0.1, σ1=0.05, σ2=0.05
    (A3)[10] ˉr1=0.1, ˉr2=0.3, β1=0.3, β2=0.0001, α1=0.0001, α2=0.3, ξ1=1, ξ2=1, δ1=2
    δ2=2, γ1=0.2, γ2=0.2, ω1=0.1, ω2=0.1, σ1=0.05, σ2=0.05
    (A4) ˉr1=0.35, ˉr2=0.15, β1=0.3, β2=0.15, α1=0.15, α2=0.2, ξ1=0.2, ξ2=0.1, δ1=0.55
    δ2=0.2, γ1=0.3, γ2=0.25, ω1=0.1, ω2=0.1, σ1=0.05, σ2=0.05
    (A5) ˉr1=0.2, ˉr2=0.4, β1=0.1, β2=0.2, α1=0.2, α2=0.3, ξ1=0.2, ξ2=0.37, δ1=0.3
    δ2=0.75, γ1=0.2, γ2=0.4, ω1=0.1, ω2=0.1, σ1=0.05, σ2=0.05
    (A6)[26] ˉr1=0.37, ˉr2=0.37, β1=1, β2=0.1, α1=0.27, α2=2, ξ1=0.2, ξ2=0.1, δ1=0.55
    δ2=0.5, γ1=0.3, γ2=0.3, ω1=0.2, ω2=0.1, σ1=0.001, σ2=0.001

     | Show Table
    DownLoad: CSV

    Example 1. Theorem 3.1 is first tested by selecting the combination (A1), (A2), and (A3) in Table 2 as the coefficients of system (1.5). See Figure 1, Figure 2, and Figure 3.

    Figure 1.  Coefficient combination (A1).
    Figure 2.  Coefficient combination (A2).
    Figure 3.  Coefficient combination (A3).

    Remark 2. It can be seen that the death rate ri is perturbed around the given mean value, which reflects the mean reversion characteristic of the Ornstein-Uhlenbeck process.

    Comparing Figure 2 with Figure 1 and Figure 3, one can conclude that the death rate is not a determining factor in population density, and other factors such as competition intensity and environmental disturbance also have a great impact on the survival of the population.

    On the basis of Theorem 3.1, we next investigate the effect of environmental perturbations on the population by choosing the parameter combination (A1) and varying the value of the parameter σi,i=1,2, in it. Let σi = 0.01, 0.08, 0.12, respectively. According to Figure 4, the population becomes more unstable as the environmental noise increases, which implies that the environmental perturbation has a great impact on the survival of the population.

    Figure 4.  The effect of different σi values on the populations.

    In order to verify the existence and uniqueness of the solution of system (1.5) more comprehensively and clearly, we perform 100 simulations of Theorem 3.1. The solid green lines in (a), (b), (c), and (d) of Figure 5 represent the average of the 100 simulated paths, and all 100 paths are represented by gray lines. It can be seen that different coefficient combinations have different solutions, and the solutions are all existing and unique. Thus, the conclusion of Theorem 3.1 can be verified.

    Figure 5.  100 path simulation figures.

    Example 2. Now we will verify the conclusion of Theorem 3.2. We still choose the combination (A1), (A2), and (A3) in Table 2 as the coefficients of system (1.5). Let q = 2, then we have E[|x(t),y(t)|2]K(2). It can be seen from Figure 6 that the expected value of the above three coefficient combinations is less than an upper bound K(q) and this upper limit is not infinite, which indicates that the two-moment of the population is bounded. See Figure 6.

    Figure 6.  Moment estimation.

    Example 3. We will now verify the conclusion of Theorem 3.3 by numerical simulation. By Table 2, We still use combination (A1), (A2) and (A3) for verification. See Figure 7

    Figure 7.  Stochastic ultimate boundedness.

    Remark 3. From the biological point of view, since the environmental resources are limited, no biological population can grow indefinitely, so we hope that the system solution is ultimately bounded. It can be seen from Figure 7 that, with the increase of time t, the probability P{x2(s)+y2(s)H} is gradually stable and greater than a constant, which means lim suptP{x2(s)+y2(s)H}1ε. Therefore, the above simulation verifies the stochastic ultimate boundedness.

    In order to further verify the conclusions of Theorem 3.2 and Theorem 3.3, we choose 100 paths for simulation as in Theorem 3.1. The left figure is the simulation result of Theorem 3.2, and the right figure is the simulation result of Theorem 3.3. See Figure 8.

    Figure 8.  100 path simulation figures.

    Example 4. We now verify the conclusion of Theorem 3.4 in terms of parameter combinations (A4), (A5), and (A6).

    In Figure 9 (a), we choose combination (A4) as the biological parameter values of system (1.5). Direct calculations shows that G1=0.343<0, it is easy to see that the parameters satisfy the condition of Theorem 3.4, and x is extinct. Figure 9 (a) confirms these.

    Figure 9.  Population extinction.

    In Figure 9 (b), we choose combination (A5) as the biological parameter values of system (1.5). Direct calculations shows that G2=0.417<0, it is easy to see that the parameters satisfy the condition of Theorem 3.4, and y is extinct. Figure 9 (b) confirms these.

    In Figure 9 (c), we choose combination (A6) as the biological parameter values of system (1.5). Direct calculations shows that G1=0.343<0 and G2=0.527<0, it is easy to see that the parameters satisfy the condition of Theorem 3.4, and all the species tend to be extinct. Figure 9 (c) confirms these.

    Now explore the effect of different G values on the population.

    In Figure 10, we select parameter combination (A4) and change the value of γ1 in it. Let γ1=0.2,0.3,0.4, respectively. It can be seen that, as the value of G1 decreases, the extinction time of population x is advanced. It can also be seen that, when the jump noise coefficient γ1<0, then the smaller γ1 is, the greater the negative impact on the population.

    Figure 10.  Effect of G1 on population x.

    In Figure 11, we select parameter combination (A5) and change the value of ˉr2 in it. Let ˉr2=0.36,0.4,0.46, respectively. It can be seen that, as the value of G2 decreases, the extinction time of population y is advanced. It can also be seen that the death rate accelerates population extinction, which is consistent with the phenomenon in nature.

    Figure 11.  Effect of G2 on population y.

    In this paper, we first introduce a stochastic two-species Schoener's competitive model. Moreover, since populations in nature are often subject by sudden random perturbations, we introduced Lévy jumps to model this phenomenon. Previous research endeavors have traditionally employed white noise or telegraph noise to simulate environmental perturbations. Nonetheless, E.Allen[27] has identified certain conceptual and practical limitations of linear functions of Gaussian white noise. In addition, the role of the Ornstein-Uhlenbeck process in population dynamics models has received relatively little attention. It has been theoretically demonstrated that the mean-reverting Ornstein-Uhlenbeck process offers more stable environmental variability than linear and nonlinear perturbations. Therefore, we incorporate the Ornstein-Uhlenbeck process into the deterministic system, thereby obtaining a stochastic two-species Schoener's competitive model with Ornstein-Uhlenbeck process. It should be noted that the relevant dynamical properties of system (1.5) with the introduction of the Ornstein-Uhlenbeck process are changed. For example, the existence and uniqueness of the global positive solution studied in the traditional model becomes the existence and uniqueness of the global solution after adding the Ornstein-Uhlenbeck process. The Lyapunov functions used in the proofs of other properties (moment boundedness, extinction) are also different.

    To the best of our ability, we prove the existence and uniqueness of the global solution, moment estimation, stochastic ultimate boundedness, and give sufficient conditions for population extinction. At present, few papers add Lévy jump or the Ornstein-Uhlenbeck process to Schoener's competitive model, and research on Schoener's competitive model mainly focus on periodic solution, stationarity, and persistence. Therefore, the study of other properties of system (1.5) has certain value.

    In fact, our model still has areas for improvement. First, since the model we studied is two-dimensional, this leads to limitations in simulating the dynamical behaviour of populations in nature. Therefore, in the future, we will expand the model from two-dimensional to n-dimensional. In addition, it is well known that, in many natural ecosystems, there is a time lag in the interaction between populations, but we have not considered the effect of time lag on the model. In the future, we will utilize the relevant theories and methods of time-lagged generalized differential equations and impulse differential equations to build the corresponding dynamical models.

    The authors declare they have not used Artificial Intelligence (AI) tools in the creation of this article.

    This study was supported by the National Natural Science Foundation of China under Grant (No.11401085), the Central University Basic Research Grant (2572021DJ04), the Heilongjiang Postdoctoral Grant (LBH-Q21059), and the Innovation and Entrepreneurship Training Program for University Students (S202310225019).

    Thanks to all anonymous reviewers for their valuable comments and helpful suggestions that made the article better.

    The authors declare there is no conflicts of interest.



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